Mystery: why is a quarter rest missing for no reason?

See screenshot, bottom stave, m49. Why is that first quarter rest missing? I’ve tried everything to get it back:

I even deleted the contents of the entire measure and attempted to rewrite it; the first quarter rest is still missing!

Screenshot above taken in Write mode, but it looks the same in Engrave mode too. I don’t know where to look to fix it.

Can you cut down the project to the problematic bars and attach it here?

I’d say the problem comes from earlier than this bar. Locate the last note before that C, check its properties and uncheck “ends voice” property.

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