Halp my midi tracks are transposing themselves

Alright…I’m just doing a simple piano roll composition here with some simple vsts, and 3 of my four vst instruments are suddenly transposing themselves a minor third down.
It’s driving me crazy. I enter in one note into the piano roll, and it comes out a minor third off. I’ve tried creating new tracks and copying the piano roll information between them. But it’s still the same. One track remains untransposed. I did nothing in the midi transpose settings. Why is this happening??!?!
Cubase 8.5

Oh man I figured it out. There are so many tiny things going on.

How did you fix it, just in case someone finds your thread on Google?